BMS107 Chapter 19 Outline
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STIMULI AND RECEPTORS
- general sense receptors - are housed in the skin, musculoskeletal organs, and viscera
- special senses: vision, hearing, equilibrium, olfaction, gustation
- receptors act as transducers that transform the energy in a stimulus into electrical (nerve) signals
VISION
- accessory structures
- conjunctiva (mucous membrane) - ocular conjunctiva, palpebral conjunctiva
- lacrimal apparatus
- lacrimal gland - produces lacrimal fluid
- nasolacrimal duct - receives fluid; fluid drains to nasal cavity
- fibrous tunic
- cornea - avascular, convex, refracts (bends) light rays
- sclera - dense irregular connective tissue, provides shape and protection
(skleros = hard)
- vascular tunic ("uvea")
- choroid - cells are filled with melanin to absorb extraneous light
- ciliary body
- in anterior portion of vascular tunic
- ciliary muscle - four bands of smooth muscle organized into a ring
- ciliary processes - extensive folds of ciliary body
- aqueous humor - fluid produced by epithelium of ciliary body
- suspensory ligaments extend from ciliary processes
and attach to lens capsule
- lens is strong yet deformable
- ciliary body and suspensory ligaments determine shape and
position of lens
- for far vision
- ciliary muscle relaxes
- tension on suspensory ligaments increases
- causes lens to flatten ("default" position)
- accommodation for near vision
- ciliary muscle contracts
- ciliary body moves anteriorly
- reduces tension in suspensory ligaments
- lens becomes more spherical and
the degree of refraction increases
- iris
- continuous with ciliary body
- pupil - hole in center of iris
- iris contains pigment-forming cells and two groups of smooth muscle fibers
- sphincter pupillae muscle (pupillary constrictor) -
arranged in concentric circles; innervated by parasympathetic division of ANS
- dilator pupillae muscle (pupillary dilator) - organized in radial pattern;
controlled by sympathetic division of ANS
- neural tunic (retina)
- pigmented layer
- neural layer - contains photoreceptors
- ora serrata - jagged margin of retina
- organization of neural retina (know locations & sequence of layers)
- photoreceptor cells
- rods - more than 100M per eye; function in dim light;
relatively high degree of convergence with bipolar cells
- cones (require high intensity light)
- low degree of convergence with bipolar cells provides visual acuity
- color vision - based on three types of cones with varying sensitivities
("blue cones," "green cones," "red cones")
- macula lutea - "yellow spot"
- fovea centralis - "central pit" - on visual axis;
contains high proportion of cones (almost no rods)
- bipolar cells - form synapses with bipolar cells
- ganglion cells - axons from ganglionic cells extend through the optic disc
and form optic nerve (no photoreceptors at optic disc)
VISUAL PATHWAY
- retina
- optic nerves
- optic chiasm - medial axons cross
(Figs. 15.8, 17.11, 19.18)
- optic tracts
- in brain, optic tract axons extend to
- thalamus (lateral geniculate nucleus)
- midbrain (for coordination of eye, head, and neck movements)
- projection fibers (optic radiations) project from thalamus to occipital lobes
CAVITIES OF THE EYE
- anterior cavity
- posterior to cornea / anterior to lens
- chambers
- anterior chamber - between iris and cornea
- posterior chamber - between iris and lens
- contains aqueous humor
- secreted into posterior chamber; flows through pupil into anterior chamber
- maintains chemical environment & creates intraocular pressure
- continually reabsorbed into scleral venous sinus then
conducted to veins
- posterior cavity
- posterior to lens / anterior to retina
- occupied by transparent, gelatinous vitreous body (between lens and retina)
- vitreous body helps maintain eye shape and supports retina
EXTERNAL EAR
- auricle - directs sound waves into...
- external auditory canal, which terminates at...
- tympanic membrane ("eardrum")
- partition between external and middle ear
- medial surface is lined by a mucosa
- vibrates in response to sound waves
MIDDLE EAR
- contains air-filled tympanic cavity
- auditory tube (pharyngotympanic tube, Eustachian tube)
- opens into nasopharynx from middle ear
- air can move through tube for pressure equalization between tympanic cavity and atmosphere
- auditory ossicles
- malleus - attached to medial surface of tympanic membrane
- incus
- stapes - footplate fits into oval window (lateral wall of inner ear)
- ossicles vibrate along with tympanic membrane
- movement of stapes initiates pressure waves in fluid within inner ear
- tympanic membrane is approximately twenty times greater in diameter than the footplate of
the stapes (concentrates force of vibrations)
- stapedius & tensor tympani - skeletal muscles;
restrict ossicle movement when loud noises occur (protect inner ear)
INNER EAR
- membranous labyrinth - fits within bony labyrinth (in petrous portion of temporal bone)
- perilymph - fluid between membranous labyrinth and wall of bony labyrinth
- endolymph - fluid within membranous labyrinth
- regions of bony labyrinth / membranous labyrinth
- vestibule (of bony lab.) - contains utricle and saccule (of memb. lab.)
- semicircular canals (of bony lab.) - contain semicircular ducts (of memb. lab.)
- cochlea (of bony lab.) - contains cochlear duct (of memb. lab.)
- vestibular complex = vestibule + semicircular canals
- hair cells
- receptors for both hearing and equilibrium
- stimulated by mechanical forces (stretch/compression/distortion)
- hearing
- cochlear duct - runs through cochlea
- basilar membrane - floor of cochlear duct
- spiral organ - consisting of hair cells and supporting cells
- hair cells extend stereocilia into tectorial membrane
- movement of basilar membrane distorts hair cells
(leads to stimulation of sensory neurons in cochlear nerve)
- auditory pathway
- cochlear nerve axons terminate in cochlear nuclei in brainstem
- secondary neurons project to thalamus, midbrain, etc.
- some axons extend from thalamus to auditory cortex
- equilibrium
- receptors in utricle and saccule detect position/acceleration of head
- tilt of head > otolithic membrane shifts >
distortion of hair cells stimulates sensory neurons
- receptors in semicircular canals detect rotation of the head
- rotation of head > endolymph moves / pushes against cupula >
distortion of hair cells stimulates sensory neurons
- vestibular sensation pathways
- vestibular branch of CN VIII
- vestibular nerve axons project to vestibular nuclei in brainstem
- vestibular nuclei integrate signals and send impulses to cerebellum,
thalamus, spinal cord, other brainstem nuclei, etc.
Chapter 19 Questions
In Textbook, Second Edition
- What Did You Learn? - 10, 11, 13, 14
- What Do You Think? - 4
- Matching - 1, 2, 3, 5, 6, 8
- Multiple Choice - 4, 6
- Content Review - 5, 6, 7, 8, 9
At Publisher's Web Site
Simple Multiple Choice - 2, 3, 4, 5, 6, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24, 27, 28, 29, 30
Difficult Multiple Choice - 1, 10, 20, 21, 22
Challenge Yourself - 1, 2, 3, 4, 6, 8, 9, 10
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Spring 2009